Do you want to contribute to improving human health?

To be a doctoral student means to devote oneself to a research project under supervision of experienced researchers and following an individual study plan. For a doctoral degree, the equivalent of four years of full-time doctoral education is required.

The research group

Fuzik Lab is looking for an applicant for a PhD position (Circuit and Systems Neuroscience) at the Dept. of Neuroscience, Karolinska Institute, Stockholm, Sweden. The PhD program is funded by the KI-NIH program in Neuroscience. The PhD student will spend 50% (year 1-2) at the Dept. of Neuroscience, Karolinska Institute and 50% (year 3-4) at the National Institutes of Health, NICHD / DIR, Bethesda, USA.

The doctoral student project and the duties of the doctoral student

The applicant will join an active, enthusiastic, and well-equipped group studying fundamental properties of periaqueductal and hypothalamic neuronal pathways and neuromodulation primarily in panic and anxiety including the aspect of sex differences.

We study both physiological and pathological states of periaqueductal neuronal populations with viral tracing, Patch-clamp electrophysiology, Patch-Seq (Fuzik et al., 2016, Nature Biotechnology, Fuzik et al. 2019, PNAS), ex vivo all-optical voltage imaging and Voltage-Seq (Csillag et al. 2023, Nature Methods), in vivo optogenetics, and add a translational angle using systems neuroscience approaches (behavior tests, head-fixed in vivo electrophysiology (Calvigioni, Fuzik et al. 2023, Nature Neuroscience).

A PhD candidate with a neuroscience background is essential. Basic (even not documented) expertise in programming (Python, Matlab) is preferable, documented experience in microscopy optics is preferred, some experience with behavior tests, optogenetics is preferred, experience in electrophysiology and/or imaging techniques will be highly preferred.

What do we offer?

A creative and inspiring environment full of expertise and curiosity. Karolinska Institutet is one of the world's leading medical universities. Our vision is to pursue the development of knowledge about life and to promote a better health for all. At Karolinska Institutet, we conduct successful medical research and hold the largest range of medical education in Sweden. As a doctoral student you are offered an individual research project, a well-educated supervisor, a vast range of elective courses and the opportunity to work in a leading research group. Karolinska Institutet collaborates with prominent universities from all around the world, which ensures opportunities for international exchanges. You will be employed on a doctoral studentship which means that you receive a contractual salary. Employees also have access to our modern gym for free and receive reimbursements for medical care.

Eligibility requirements for doctoral education

In order to participate in the selection for a doctoral position, you must meet the following general (A) and specific (B) eligibility requirements at latest by the application deadline.

It is your responsibility to certify eligibility by following the instructions on the web page Entry requirements (eligibility) for doctoral education.

A) General eligibility requirement

You meet the general eligibility requirement for doctoral/third-cycle/PhD education if you:

  1. have been awarded a second-cycle/advanced/master qualification (i.e. master degree), or
  2. have satisfied the requirements for courses comprising at least 240 credits of which at least 60 credits were awarded in the advanced/second-cycle/master level, or
  3. have acquired substantially equivalent knowledge in some other way in Sweden or abroad.*

Follow the instructions on the web page Entry requirements (eligibility) for doctoral education.

*If you claim equivalent knowledge, follow the instructions on the web page Assessing equivalent knowledge for general eligibility for doctoral education.


B) Specific eligibility requirement

You meet the specific eligibility requirement for doctoral/third-cycle/PhD education if you:

- Show proficiency in English equivalent to the course English B/English 6 at Swedish upper secondary school.

Follow the instructions on the web page English language requirements for doctoral education.

Verification of your documents Karolinska Institutet checks the authenticity of your documents. Karolinska Institutet reserves the right to revoke admission if supporting documents are discovered to be fraudulent. Submission of false documents is a violation of Swedish law and is considered grounds for legal action.

(A) and (B) can only be certified by the documentation requirement for doctoral education.

Skills and personal qualities

[Enter here skills and personal qualities necessary or desirable/advantageous for this project and for the ability to reach the outcomes of the doctoral education. Make it clear for the applicants what is necessary and what is only desirable. It is important that this section is well thought-through in order to help in the selection process.]

Terms and conditions

The doctoral student will be employed on a doctoral studentship maximum 4 years full-time.

Application process

Submit your application and supporting documents through the Varbi recruitment system. Use the button in the top right corner and follow the instructions. We prefer that your application is written in English, but you can also apply in Swedish.

Your application must contain the following documents:

- A personal letter and a curriculum vitae
- Degree projects and previous publications, if any
- Any other documentation showing the desirable skills and personal qualities described above
- Documents certifying your general eligibility (see A above)
- Documents certifying your specific eligibility (see B above)


A selection will be made among eligible applicants on the basis of the ability to benefit from doctoral education. The qualifications of the applicants will be evaluated on an overall basis.

Karolinska Institutet uses the following bases of assessment:

- Documented subject knowledge of relevance to the area of research
- Analytical skill
- Other documented knowledge or experience that may be relevant to doctoral studies in the subject.

All applicants will be informed when the recruitment is completed.


Want to make a difference? Join us and contribute to better health for all

Type of employment PhD placement
Contract type Full time
First day of employment As soon as possible
Salary Monthly salary
Number of positions 1
Full-time equivalent 100
City Solna
County Stockholms län
Country Sweden
Reference number STÖD 2-641/2024
  • Janos Fuzik,
Published 08.Feb.2024
Last application date 07.Mar.2024 11:59 PM CET

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